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Useful Temperate Plants

Pinus halepensis

Mill.

Pinaceae

+ Synonyms

Pinus hierosolimitana Duhamel

Pinus maritima Mill.

Pinus maritima Aiton

Pinus penicillus Lapeyr.

Pinus arabica Sieber ex Spreng.

Pinus genuensis J.Cook

Pinus hispanica J.Cook

Pinus carica D.Don

Pinus abásica Carrière

Pinus abchasica Carrière

Pinus loiseleuriana Carrière

Pinus paroliniana Webb ex Carrière

Pinus pseudohalepensis Denhardt ex Carrière

Pinus parolinii Vis.

Pinus colchica Booth ex Gordon & Glend.

Pinus saportae Rouy in G.Rouy & J.Foucaud

Pinus ceciliae Llorens & L.Llorens

Common Name: Aleppo Pine

No Image.

General Information

Pinus halepensis is an evergreen tree with an irregular, rounded crown; it can grow 15 - 25 metres tall. The crown can become very wind-shaped, especially when growing near the sea. The bole can be up to 150cm in diameter[
329
Title
The Gymnosperm Database
Publication
 
Author
 
Website
http://www.conifers.org/index.htm
Publisher
 
Year
0
ISBN
 
Description
A vast amount of information on conifers and families of gymnosperms.
].
The trees are often harvested from the wild for their resin, which is traded, and as a source of firewood for local use. The wood is also sometimes used, and the plant has sometimes been cultivated in plantations.
Pinus halepensis has a very extensive extent of occurrence and although it is considered to be threatened locally (Algarve in Portugal, Costa Brava in Spain) elsewhere it is stable or perhaps expanding as its economic use for timber has at least in its native habitat diminished. The plant is classified as 'Least Concern' in the IUCN Red List of Threatened Species(2013)[
338
Title
IUCN Red List of Threatened Species
Publication
 
Author
 
Website
http://www.iucnredlist.org/
Publisher
 
Year
0
ISBN
 
Description
A list of plants under threat and facing possible extinction, usually with brief details of the threats and information on habitat.
].
Ill considered plantations established in the southern hemisphere have given rise to invasiveness of this species; in Australia, New Zealand, and South Africa. Pinus halepensis is now a serious weed threatening natural and often species-rich vegetation as well as land used for agriculture[
338
Title
IUCN Red List of Threatened Species
Publication
 
Author
 
Website
http://www.iucnredlist.org/
Publisher
 
Year
0
ISBN
 
Description
A list of plants under threat and facing possible extinction, usually with brief details of the threats and information on habitat.
].

Known Hazards

The wood, sawdust and resins from various species of pine can cause dermatitis in sensitive people[
222
Title
A Field Guide to Medicinal Plants. Eastern and Central N. America.
Publication
 
Author
Foster. S. & Duke. J. A.
Publisher
Houghton Mifflin Co.
Year
1990
ISBN
0395467225
Description
A concise book dealing with almost 500 species. A line drawing of each plant is included plus colour photographs of about 100 species. Very good as a field guide, it only gives brief details about the plants medicinal properties.
].

Botanical References

11
Title
Trees and Shrubs Hardy in Great Britain. Vol 1 - 4 and Supplement.
Publication
 
Author
Bean. W.
Publisher
Murray
Year
1981
ISBN
-
Description
A classic with a wealth of information on the plants, but poor on pictures.
,
50
Title
Flora Europaea
Publication
 
Author
?
Publisher
Cambridge University Press
Year
1964
ISBN
-
Description
An immense work in 6 volumes (including the index). The standard reference flora for Europe, it is very terse though and with very little extra information. Not for the casual reader.
,
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.

Range

Throughout most of the Mediterranean region, but absent from Egypt.

Habitat

Sometimes found in dense woodland, but more commonly scattered in maquis or garrigue vegetation on sunny hills and slopes down to the sea shore, most commonly on limestone and dolomite; growing at elevations from sea level to 1,700 metres[
338
Title
IUCN Red List of Threatened Species
Publication
 
Author
 
Website
http://www.iucnredlist.org/
Publisher
 
Year
0
ISBN
 
Description
A list of plants under threat and facing possible extinction, usually with brief details of the threats and information on habitat.
].

Properties

Weed PotentialYes
Conservation StatusLeast Concern
Edibility Rating *  *
Medicinal Rating *  *
Other Uses Rating *  *  *
HabitEvergreen Tree
Height15.00 m
PollinatorsWind
Self-fertileNo
Cultivation StatusCultivated, Wild

Cultivation Details

Pinus halepensis is native to the Mediterranean region, where the summers are hot and dry and the winters cool and moist. It is found in the hotter parts of the region, where brush and forest fires are frequent[
338
Title
IUCN Red List of Threatened Species
Publication
 
Author
 
Website
http://www.iucnredlist.org/
Publisher
 
Year
0
ISBN
 
Description
A list of plants under threat and facing possible extinction, usually with brief details of the threats and information on habitat.
]. The plant is not very cold-hardy, being able to tolerate short periods with temperatures down to around -10°c when fully dormant[
329
Title
The Gymnosperm Database
Publication
 
Author
 
Website
http://www.conifers.org/index.htm
Publisher
 
Year
0
ISBN
 
Description
A vast amount of information on conifers and families of gymnosperms.
]. Plants become more hardy with age, some older specimens in southern Britain have been known to withstand temperatures down to -17°c without damage[
11
Title
Trees and Shrubs Hardy in Great Britain. Vol 1 - 4 and Supplement.
Publication
 
Author
Bean. W.
Publisher
Murray
Year
1981
ISBN
-
Description
A classic with a wealth of information on the plants, but poor on pictures.
].
Pine species in general require a sunny position if they are to grow well and, whilst the different species have their own soil preferences, most will thrive in a light well-drained sandy or gravelly loam[
1
Title
RHS Dictionary of Plants plus Supplement. 1956
Publication
 
Author
F. Chittendon.
Publisher
Oxford University Press
Year
1951
ISBN
-
Description
Comprehensive listing of species and how to grow them. Somewhat outdated, it has been replaced in 1992 by a new dictionary (see [200]).
,
11
Title
Trees and Shrubs Hardy in Great Britain. Vol 1 - 4 and Supplement.
Publication
 
Author
Bean. W.
Publisher
Murray
Year
1981
ISBN
-
Description
A classic with a wealth of information on the plants, but poor on pictures.
]. Dislikes poorly drained moorland soils[
1
Title
RHS Dictionary of Plants plus Supplement. 1956
Publication
 
Author
F. Chittendon.
Publisher
Oxford University Press
Year
1951
ISBN
-
Description
Comprehensive listing of species and how to grow them. Somewhat outdated, it has been replaced in 1992 by a new dictionary (see [200]).
]. Established plants are very tolerant of drought[
81
Title
Conifers.
Publication
 
Author
Rushforth. K.
Publisher
Christopher Helm
Year
1987
ISBN
0-7470-2801-X
Description
Deals with conifers that can be grown outdoors in Britain. Good notes on cultivation and a few bits about plant uses.
,
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
], they succeed in poor chalky soils and also in poor dry sandy soils[
166
Title
The Milder Garden.
Publication
 
Author
Taylor. J.
Publisher
Dent
Year
1990
ISBN
 
Description
A good book on plants that you didn't know could be grown outdoors in Britain.
]. Fairly wind-resistant[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
].
An average size tree can produce 3 - 4 kilograms of resin per year[
329
Title
The Gymnosperm Database
Publication
 
Author
 
Website
http://www.conifers.org/index.htm
Publisher
 
Year
0
ISBN
 
Description
A vast amount of information on conifers and families of gymnosperms.
].
The seed cones are 6 - 12cm x 4 - 7cm. They take 3 years to mature and can remain on the branches for several years thereafter without opening. The cones usually only open in very strong summer heat or as the result of the heat from a forest fire[
329
Title
The Gymnosperm Database
Publication
 
Author
 
Website
http://www.conifers.org/index.htm
Publisher
 
Year
0
ISBN
 
Description
A vast amount of information on conifers and families of gymnosperms.
,
338
Title
IUCN Red List of Threatened Species
Publication
 
Author
 
Website
http://www.iucnredlist.org/
Publisher
 
Year
0
ISBN
 
Description
A list of plants under threat and facing possible extinction, usually with brief details of the threats and information on habitat.
].
Plants are strongly outbreeding, self-fertilized seed usually grows poorly[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
]. They hybridize freely with other members of this genus[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
].
Leaf secretions inhibit the germination of seeds, thereby reducing the amount of pants that can grow under the tree[
18
Title
Companion Plants.
Publication
 
Author
Philbrick H. and Gregg R. B.
Publisher
Watkins
Year
1979
ISBN
-
Description
Details of beneficial and antagonistic relationships between neighbouring plants.
].
Plants in this genus are notably susceptible to honey fungus[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
].

Edible Uses

A resin from the trunk is used for chewing and for flavouring wine[
177
Title
Plants for Human Consumption
Publication
 
Author
Kunkel. G.
Publisher
Koeltz Scientific Books
Year
1984
ISBN
3874292169
Description
An excellent book for the dedicated. A comprehensive listing of Latin names with a brief list of edible parts.
,
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
].

A vanillin flavouring is obtained as a by-product of other resins that are released from the pulpwood[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
].

The seeds of all Pinus species are more or less edible, and some are of good size and make very tasty and nutritious foods, often eaten in quantitiy. Others can be less desireable, either having a strongly resinous flavour, being bitter or, more commonly, rather too small and fiddly to make it very worthwhle even trying to eat them,
We have no specific information of the desireability of this species, though they are rather small (the size given includes the shell)[
K
Title
Plants for a Future
Author
Ken Fern
Description
Notes from observations, tasting etc at Plants For A Future and on field trips.
].
Seed 5-6 mm long with a 2.5 cm wing[
329
Title
The Gymnosperm Database
Publication
 
Author
 
Website
http://www.conifers.org/index.htm
Publisher
 
Year
0
ISBN
 
Description
A vast amount of information on conifers and families of gymnosperms.
].

Medicinal

Oil of turpentine, obtained from the oleo-resin of all pine trees, is antiseptic, antispasmodic, astringent, diuretic, rubefacient, stimulant and vermifuge[
4
Title
A Modern Herbal.
Publication
 
Author
Grieve.
Publisher
Penguin
Year
1984
ISBN
0-14-046-440-9
Description
Not so modern (1930's?) but lots of information, mainly temperate plants.
,
329
Title
The Gymnosperm Database
Publication
 
Author
 
Website
http://www.conifers.org/index.htm
Publisher
 
Year
0
ISBN
 
Description
A vast amount of information on conifers and families of gymnosperms.
]. It is a valuable remedy used internally in the treatment of kidney and bladder complaints and is used both internally and as a rub and steam bath in the treatment of rheumatic affections[
4
Title
A Modern Herbal.
Publication
 
Author
Grieve.
Publisher
Penguin
Year
1984
ISBN
0-14-046-440-9
Description
Not so modern (1930's?) but lots of information, mainly temperate plants.
]. It is also very beneficial to the respiratory system and so is useful in treating diseases of the mucous membranes and respiratory complaints such as coughs, colds, influenza and TB[
4
Title
A Modern Herbal.
Publication
 
Author
Grieve.
Publisher
Penguin
Year
1984
ISBN
0-14-046-440-9
Description
Not so modern (1930's?) but lots of information, mainly temperate plants.
].
Applied externally in the form of liniment plasters, poultices, herbal steam baths and inhalers, it is a very beneficial treatment for a variety of skin complaints, wounds, sores, burns, boils etc[
4
Title
A Modern Herbal.
Publication
 
Author
Grieve.
Publisher
Penguin
Year
1984
ISBN
0-14-046-440-9
Description
Not so modern (1930's?) but lots of information, mainly temperate plants.
].

Agroforestry Uses:

The needles contain a substance called terpene, this is released when rain washes over the needles and it has a negative effect on the germination of some plants, including wheat[
201
Title
A - Z of Companion Planting.
Publication
 
Author
Allardice.P.
Publisher
Cassell Publishers Ltd.
Year
1993
ISBN
0-304-34324-2
Description
A well produced and very readable book.
].
A fairly wind-tolerant tree, it can be used in shelterbelt plantings[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
].
Trees have an extensive root system and they are planted on sand dunes in order to stabilize them[
81
Title
Conifers.
Publication
 
Author
Rushforth. K.
Publisher
Christopher Helm
Year
1987
ISBN
0-7470-2801-X
Description
Deals with conifers that can be grown outdoors in Britain. Good notes on cultivation and a few bits about plant uses.
].

Other Uses

A tan or green dye is obtained from the needles[
168
Title
Nature's Colors - Dyes from Plants.
Publication
 
Author
Grae. I.
Publisher
MacMillan Publishing Co. New York.
Year
1974
ISBN
0-02-544950-8
Description
A very good and readable book on dyeing.
].

Tannin is obtained from the bark[
100
Title
Flowers of Europe - A Field Guide.
Publication
 
Author
Polunin. O.
Publisher
Oxford University Press
Year
1969
ISBN
0192176218
Description
An excellent and well illustrated pocket guide for those with very large pockets. Also gives some details on plant uses.
].

A volatile oil obtained from the branches and leaves of the plant is used as an ingredient in commercial cosmetic preparations as a masking agent[
1243
Title
CosIng
Publication
 
Author
 
Website
http://ec.europa.eu/growth/tools-databases/cosing/
Publisher
 
Year
0
ISBN
 
Description
CosIng is the European Commission database for information on cosmetic substances and ingredients.
].
An extract of the needles is used as an ingredient in commercial cosmetic preparations as an antimicrobial, antioxidant and antiseborrhoeic[
1243
Title
CosIng
Publication
 
Author
 
Website
http://ec.europa.eu/growth/tools-databases/cosing/
Publisher
 
Year
0
ISBN
 
Description
CosIng is the European Commission database for information on cosmetic substances and ingredients.
].[
1243
Title
CosIng
Publication
 
Author
 
Website
http://ec.europa.eu/growth/tools-databases/cosing/
Publisher
 
Year
0
ISBN
 
Description
CosIng is the European Commission database for information on cosmetic substances and ingredients.
].
The seed oil is used as an ingredient in commercial cosmetic preparations as an emollient and skin conditioner[
1243
Title
CosIng
Publication
 
Author
 
Website
http://ec.europa.eu/growth/tools-databases/cosing/
Publisher
 
Year
0
ISBN
 
Description
CosIng is the European Commission database for information on cosmetic substances and ingredients.
].

The wood is rich in resin and is still tapped locally for that product; plantations have been established for this use in Greece[
338
Title
IUCN Red List of Threatened Species
Publication
 
Author
 
Website
http://www.iucnredlist.org/
Publisher
 
Year
0
ISBN
 
Description
A list of plants under threat and facing possible extinction, usually with brief details of the threats and information on habitat.
].
'Greek turpentine' is obtained from the stems[
46
Title
Dictionary of Economic Plants.
Publication
 
Author
Uphof. J. C. Th.
Publisher
Weinheim
Year
1959
ISBN
-
Description
An excellent and very comprehensive guide but it only gives very short descriptions of the uses without any details of how to utilize the plants. Not for the casual reader.
,
61
Title
A Dictionary of Plants Used by Man.
Publication
 
Author
Usher. G.
Publisher
Constable
Year
1974
ISBN
0094579202
Description
Forget the sexist title, this is one of the best books on the subject. Lists a very extensive range of useful plants from around the world with very brief details of the uses. Not for the casual reader.
,
171
Title
Economic Botany.
Publication
 
Author
Hill. A. F.
Publisher
The Maple Press
Year
1952
ISBN
-
Description
Not very comprehensive, but it is quite readable and goes into some detail about the plants it does cover.
]. The resins are obtained by tapping the trunk, or by destructive distillation of the wood[
4
Title
A Modern Herbal.
Publication
 
Author
Grieve.
Publisher
Penguin
Year
1984
ISBN
0-14-046-440-9
Description
Not so modern (1930's?) but lots of information, mainly temperate plants.
,
64
Title
Vegetable Gums and Resins.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
 
ISBN
81-88818-17-8
Description
A very good book dealing with the subject in a readable way.
]. In general, trees from warmer areas of distribution give the higher yields[
64
Title
Vegetable Gums and Resins.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
 
ISBN
81-88818-17-8
Description
A very good book dealing with the subject in a readable way.
]. Oil of Turpentine consists of an average of 20% of the oleo-resin[
64
Title
Vegetable Gums and Resins.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
 
ISBN
81-88818-17-8
Description
A very good book dealing with the subject in a readable way.
] and is separated by distillation[
4
Title
A Modern Herbal.
Publication
 
Author
Grieve.
Publisher
Penguin
Year
1984
ISBN
0-14-046-440-9
Description
Not so modern (1930's?) but lots of information, mainly temperate plants.
,
64
Title
Vegetable Gums and Resins.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
 
ISBN
81-88818-17-8
Description
A very good book dealing with the subject in a readable way.
]. Turpentine has a wide range of uses including as a solvent for waxes etc, for thinning and dissolving varnish and paints, a shoe polish, medicinal etc[
4
Title
A Modern Herbal.
Publication
 
Author
Grieve.
Publisher
Penguin
Year
1984
ISBN
0-14-046-440-9
Description
Not so modern (1930's?) but lots of information, mainly temperate plants.
,
329
Title
The Gymnosperm Database
Publication
 
Author
 
Website
http://www.conifers.org/index.htm
Publisher
 
Year
0
ISBN
 
Description
A vast amount of information on conifers and families of gymnosperms.
].
Rosin is also obtained by distillation of pine resin. This is used in paper glue and soap manufacturing, as a constituent of varnishes and paints, and for coating the bows of stringed musical instruments such as violins[
4
Title
A Modern Herbal.
Publication
 
Author
Grieve.
Publisher
Penguin
Year
1984
ISBN
0-14-046-440-9
Description
Not so modern (1930's?) but lots of information, mainly temperate plants.
,
329
Title
The Gymnosperm Database
Publication
 
Author
 
Website
http://www.conifers.org/index.htm
Publisher
 
Year
0
ISBN
 
Description
A vast amount of information on conifers and families of gymnosperms.
].
Pitch can also be obtained from the resin and is used for waterproofing, as a wood preservative etc[
329
Title
The Gymnosperm Database
Publication
 
Author
 
Website
http://www.conifers.org/index.htm
Publisher
 
Year
0
ISBN
 
Description
A vast amount of information on conifers and families of gymnosperms.
].

Because of its small size and poor form, the wood of most trees is of low quality and is of little value as timber. It has been used for rough construction, mine props, railway sleepers, and telephone poles[
100
Title
Flowers of Europe - A Field Guide.
Publication
 
Author
Polunin. O.
Publisher
Oxford University Press
Year
1969
ISBN
0192176218
Description
An excellent and well illustrated pocket guide for those with very large pockets. Also gives some details on plant uses.
,
338
Title
IUCN Red List of Threatened Species
Publication
 
Author
 
Website
http://www.iucnredlist.org/
Publisher
 
Year
0
ISBN
 
Description
A list of plants under threat and facing possible extinction, usually with brief details of the threats and information on habitat.
].
The wood is presently used as firewood and for making charcoal[
338
Title
IUCN Red List of Threatened Species
Publication
 
Author
 
Website
http://www.iucnredlist.org/
Publisher
 
Year
0
ISBN
 
Description
A list of plants under threat and facing possible extinction, usually with brief details of the threats and information on habitat.
]. Witches brooms are common in this pine and could be a source of dwarf cultivars in Mediterranean countries; the species is generally not hardy in more northern latitudes. It has been planted as wind breaks and to stop soil erosion on dry slopes.

Propagation

It is best to sow the seed in individual pots in a cold frame as soon as it is ripe if this is possible otherwise in late winter. A short stratification of 6 weeks at 4°c can improve the germination of stored seed[
80
Title
Hardy Woody Plants from Seed.
Publication
 
Author
McMillan-Browse. P.
Publisher
Grower Books
Year
1985
ISBN
0-901361-21-6
Description
Does not deal with many species but it is very comprehensive on those that it does cover. Not for casual reading.
]. Plant seedlings out into their permanent positions as soon as possible and protect them for their first winter or two[
11
Title
Trees and Shrubs Hardy in Great Britain. Vol 1 - 4 and Supplement.
Publication
 
Author
Bean. W.
Publisher
Murray
Year
1981
ISBN
-
Description
A classic with a wealth of information on the plants, but poor on pictures.
]. Plants have a very sparse root system and the sooner they are planted into their permanent positions the better they will grow[
K
Title
Plants for a Future
Author
Ken Fern
Description
Notes from observations, tasting etc at Plants For A Future and on field trips.
]. Trees should be planted into their permanent positions when they are quite small, between 30 and 90cm[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
]. We actually plant them out when they are about 5 - 10cm tall. So long as they are given a very good weed-excluding mulch they establish very well[
K
Title
Plants for a Future
Author
Ken Fern
Description
Notes from observations, tasting etc at Plants For A Future and on field trips.
]. Larger trees will check badly and hardly put on any growth for several years. This also badly affects root development and wind resistance[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
].
Cuttings. This method only works when taken from very young trees less than 10 years old. Use single leaf fascicles with the base of the short shoot. Disbudding the shoots some weeks before taking the cuttings can help. Cuttings are normally slow to grow away[
81
Title
Conifers.
Publication
 
Author
Rushforth. K.
Publisher
Christopher Helm
Year
1987
ISBN
0-7470-2801-X
Description
Deals with conifers that can be grown outdoors in Britain. Good notes on cultivation and a few bits about plant uses.
].
Cite as: Temperate Plants Database, Ken Fern. temperate.theferns.info. 2019-09-19. <temperate.theferns.info/plant/Pinus+halepensis>

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